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In architecture, a keystone is the wedge-shaped stone placed at the apex of an arch. It is crucial because it locks the other stones, or voussoirs, into place, thereby stabilizing the entire structure.12

Beyond its literal architectural meaning, "keystone" is also used metaphorically to describe something essential that supports a larger system or concept. This could be a fundamental principle, a vital species in an ecosystem (known as a keystone species), or a foundational element upon which other things depend.123

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