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The sarcotesta is the outer, typically soft, fleshy part of the testa, which is the protective outer layer of a seed. In some seeds, like those of cycads, the sarcotesta is a prominent feature.1

In angiosperms, seeds develop from ovules and are enclosed by one or two integuments that form the seed coat, or testa. This seed coat can be hard and stony, or it may feature a fleshy sarcotesta on the outside with a stony layer (sclerotesta) inside. Other seed coverings can include wings or ornamentation, and some seeds have an additional layer called an aril, which is an outgrowth of the funiculus.2

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2 Britannica Sources
  1. sarcotesta
  2. angiosperm - Seeds

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