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Heterospory is a botanical term describing the production of two distinct types of spores: microspores and megaspores. This occurs in some vascular plants, known as pteridophytes, and all seed plants.12

Microspores develop into microgametophytes, which produce male gametes (sperm), while megaspores develop into megagametophytes, which produce female gametes (eggs). This contrasts with homospory, where only one type of spore is produced, leading to bisexual gametophytes. The evolution of heterospory, which occurred independently in several plant groups by the middle Devonian Period, is considered an advancement over homospory. It allows for obligate cross-fertilization, increasing genetic recombination and reducing mortality rates by enabling gametophytes to develop quickly within the protective spore wall.134

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