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The pupa is a life stage in insects that undergo complete metamorphosis, occurring between the larval and adult stages. During this phase, larval structures break down, and adult features, such as wings, begin to form. This transformation is regulated by hormones.

Common examples include the chrysalis of butterflies and the cocoon of moths. Within these protective coverings, the caterpillar develops into an adult. The pupa can be exarate (appendages not attached to the pupal skin), obtect (appendages attached), or coarctate (within the shed larval exoskeleton).

A feature found on many pupae, particularly in moths and butterflies, is the cremaster. This is a spiny pad or spike at the posterior end that often attaches the pupa to silk fibers spun by the larva, helping to secure it in place.12

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