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In botany, the calyptrogen is a layer of cells responsible for originating the root cap. The root cap is a protective layer of parenchyma cells that shields the root apical meristem as the root grows through the soil. New cells for the root cap are continuously added by the calyptrogen. This specialized tissue is part of the root apex, a region of active cell division that gives rise to all primary root tissues. The calyptrogen is one of the histogens, or groups of initial cells, proposed in the histogen theory of root apical organization.123

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