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An areole is a specialized structure found on cacti, from which spines, flowers, stems, or roots grow. These small, cushion-like areas are unique to the cactus family (Cactaceae) and are essentially modified branches. In most cacti, leaves are absent or reduced to spines, and the stem takes over the photosynthetic functions. The areole is where these spines, and often flowers, develop. While the term "areola" can refer to a small area or a colored ring around something (like a nipple), in plant anatomy, "areole" specifically denotes this cactus structure.123

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