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Taxonomy See also list of beetles. tortoise beetle, (subfamily Cassidinae), any member of more than 3,000 beetle species that resemble a turtle because of the forward and sideways extensions of the body. Both adults and larvae of some species are destructive to garden plants and sweet potatoes. Tortoise beetles range between 5 and 12 mm (less than 0.5 inch) in length, and the larvae are spiny. Tropical species are among the most brilliantly coloued of...