Which languages contributed loanwords to South Sakhalin Nivkh?

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Paleo-Siberian languages Distribution of Paleo-Siberian languages.

South Sakhalin Nivkh contains a considerable number of loanwords from the Ainu language, which is native to northern Japan. Additionally, during the first half of the 20th century, the language was hospitable to potential loanwords from Japanese; however, these terms never became fully acculturated because Japanese hegemony over South Sakhalin ended after 1945.1