Which South American nomads lived mainly in canoes in the upper Paraguay River marshes?

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The GuatĂł people are the South American nomads who traditionally lived in the marshes of the upper Paraguay River.12

These aquatic nomads spent much of their lives in dugout canoes, relying on fishing and hunting aquatic animals—such as cayman—for their subsistence. When not in their canoes, they constructed temporary shelters on small islands that remained slightly above the flood level. Their social structure was organized around the nuclear family, and they were divided into three local subtribes, each led by a headman.12