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Which South American Indigenous languages use tone to distinguish meaning?

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Many South American Indigenous languages utilize pitch-stress systems, where high and low tones are used on stressed syllables to distinguish meaning. These systems are very common in language families such as Panoan, Huitotoan, and Chibchan.1

While these pitch-stress systems are widespread, more complex tonal systems are considered rare in the region. Examples of these more complex systems include:

Acaricuara: Features a three-tone system.1
Mundurukú (Mundurucú): Features a four-tone system.1
Tikuna: Features a five-tone system.1

In linguistics, tone languages use relative pitch to differentiate words that are otherwise identical in their consonant and vowel sequences. While some South American languages incorporate these tonal distinctions, it is important to distinguish these linguistic features from musical scales, which also utilize varying tones but serve a different function in cultural expression.234