What does the colon symbol show in the Universal Decimal Classification?

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In the Universal Decimal Classification, the colon symbol (:) is used to indicate a relationship between two or more subjects.1

This technique of synthesis allows for the creation of complex notations by linking different parts of the classification schedules. For example, if the notation for iron and steel is 669.1 and the notation for sulfur is 546.22, the compound subject of sulfur in iron and steel is represented as 669.1:546.22. The colon is the most commonly used sign in this system to connect these distinct aspects of a subject.1