What discoveries did the Genographic Project make about Khoisan maternal lineages?

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Large illustrated genealogical tree with detailed branches and leaves, featuring family names and a central family crest, representing the Lippincott family lineage. — Genealogy Genealogical tree of the Richard and Abigail Lippincott family in America, constructed and published by Charles Lippincott, 1880.

The Genographic Project analyzed mitochondrial DNA (mtDNA) to trace maternal lineages and investigate human migration patterns. Regarding the Khoisan peoples of South Africa, researchers discovered that this group diverged from other Homo sapiens populations between 150,000 and 90,000 years ago. This finding suggests that distinct maternal lineages were already well established at a very early stage in human history.1