How many vowels did Hittite, Palaic, and Lycian have?

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Map of ancient Anatolia and northern Syria showing the geographic distribution of Anatolian languages, including Hittite, Hattian, Lydian, Phrygian, Palaic, Carian, Lycian, Pisidian, Hurrian, and Urartean. Major historical sites such as Hattusa, Kanesh, and Ugarit are marked, with the Black Sea and Mediterranean Sea bordering the region. — Anatolian languages Distribution of the Anatolian languages.

Hittite, Palaic, and Lycian each feature a four-vowel system consisting of a, e, i, and u. This system represents a reduction from the five-vowel system of Proto-Indo-European, caused by the merger of the inherited o vowel with other vowels.1

While the Lycian alphabet itself contains six vowel symbols, linguistic analysis of the Anatolian languages identifies the underlying phonological system of Lycian as having four vowels.21