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How many short and long vowels did Proto-Dravidian have?

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Thematic map of South Asia showing the distribution of Dravidian languages, with different regions shaded or marked to indicate where specific Dravidian languages and subgroups are spoken. Most languages are concentrated in southern and central India, with some northern outliers in Pakistan, Afghanistan, and eastern India. A key identifies the languages by number and subgroup, and striped areas indicate regions where several subgroups overlap. — Dravidian languages Distribution of Dravidian languages.

The Proto-Dravidian sound system consisted of five short vowels (/i/, /e/, /a/, /o/, /u/) and their five corresponding long vowels (/ī/, /ē/, /ā/, /ō/, /ū/).1