How does the cluster-of-features approach define membership in a species?

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In the philosophy of biology, the cluster-of-features approach—which has roots in the work of the French botanist Michel Adanson and was later advocated by William Whewell—defines membership in a species based on a collection of traits rather than a single defining characteristic.1

Under this framework, an organism is considered a member of a species if it possesses a sufficient number of features from a designated set. Crucially, no single feature within that cluster is considered strictly necessary for membership. This approach serves as a potential solution to the limitations of traditional morphological definitions, which often struggle to account for individuals that lack specific traits—such as rationality in humans—yet are still clearly recognized as belonging to the species.1