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Did Ardipithecus have adaptations for knuckle-walking?

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skeleton of “Ardi” — Ardipithecus Reconstructed frontal view of the skeleton of “Ardi,” a specimen belonging to the early hominid species *Ardipithecus ramidus*.

No, Ardipithecus did not have adaptations for knuckle-walking.

According to the fossil evidence from the partial skeleton known as Ardi, the hands of Ar. ramidus lacked the long metacarpals and wrist-stabilizing features characteristic of knuckle-walking apes. Instead, the wrist was capable of bending upward, and the fingers were long. Researchers have found no evidence of knuckle-walking adaptations in the arms or hands of this genus.1

While some scientific debate exists regarding whether human ancestors evolved from a knuckle-walking lineage, the specific anatomical findings for Ardipithecus indicate that it was already adapted for a combination of arboreal climbing and a form of bipedalism, rather than the knuckle-walking locomotion seen in modern African apes.12