Why can damage to outer hair cells make quiet sounds harder to hear?

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Outer hair cells play a critical role in hearing by acting as mechanical-electrical stimulators and feedback elements that modify and enhance the sensitivity of inner hair cells.1

Because inner hair cells are positioned on the bony shelf of the osseous spiral lamina rather than the basilar membrane, they are less easily stimulated by sound vibrations on their own. Outer hair cells help bridge this gap by contracting rhythmically in response to sound, which effectively amplifies the stimulus for the inner hair cells. When outer hair cells are damaged or destroyed, this amplification mechanism is lost, causing the threshold required to trigger the inner hair cells to rise significantly—often by as much as 40 dB—thereby making quiet sounds much harder to detect.1