Which marine invertebrates suffered the greatest losses during the Permian extinction?

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During the Permian extinction, shallow warm-water marine invertebrates experienced the most protracted and severe losses. The groups that suffered the greatest declines included:1

  • Trilobites: This group of arthropods made their final appearance in the fossil record at the end of the Permian.12
  • Corals: Both rugose and tabulate corals became extinct by the end of the Paleozoic Era.12
  • Echinoderms: Two major groups, the blastoids and crinoids, were hit particularly hard. Blastoids, which had persisted in some regions until the end of the period, disappeared entirely.12
  • Brachiopods: Several superfamilies, including productaceans, chonetaceans, and richthofeniaceans, vanished.2
  • Bryozoans: Cryptostomate and fenestrate groups, which were common in Paleozoic reefs, did not survive the crisis.2
  • Foraminiferans: Fusulinid foraminiferans, which served as important index fossils, also went extinct.2

Overall, these extinctions reduced marine invertebrate diversity to roughly 5 percent of their Guadalupian maxima, marking their lowest diversity levels since the end of the Ordovician Period.1