Which fossil marks the beginning of the Devonian System at the Klonk section?

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The beginning of the Devonian System at the Klonk section in the Czech Republic is marked by the first occurrence of the graptolite species Monograptus uniformis.12

This horizon, which defines the base of the Monograptus uniformis biozone, serves as the international standard—often referred to as a "golden spike"—for the Silurian-Devonian boundary. While this graptolite taxon is the primary marker, the boundary is further reinforced by associated conodont and trilobite taxa.2