What discoveries did Herophilus make through anatomical study?

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Two surgeons in scrubs and masks perform laparoscopic surgery on a patient in a brightly lit operating room, with medical equipment and a monitor displaying the procedure. — medicine Surgeons performing a laparoscopy.

Herophilus, often called the father of anatomy, made significant contributions to medical science by performing some of the first public dissections on human cadavers in Alexandria.1

His key anatomical discoveries and observations include:

Nervous System: He identified the brain as the center of the nervous system and classified nerve trunks as either motor or sensory, distinguishing them from tendons and blood vessels.1
Brain Structure: He studied the brain's ventricles and traced the sinuses of the dura mater to a junction now known as the torcular Herophili.1
Organ Descriptions: He provided detailed accounts of the liver, pancreas, salivary glands, and the genital organs. He also described and named the duodenum and the prostate gland.1
Physiology: He was the first to measure the pulse, utilizing a water clock for precision.1