How is the Permian Period divided into the Cisuralian Guadalupian and Lopingian epochs?

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The Permian Period is divided into three epochs—the Early (Cisuralian), Middle (Guadalupian), and Late (Lopingian)—which correspond to their respective rock series. These divisions are defined by the emergence of specific species of conodonts, which are primitive chordates with tooth-shaped fossil remains.1

Each epoch is further subdivided into internationally recognized ages based on these faunal successions:

  • Cisuralian Epoch: Named for the Ural Mountains, it includes the Asselian, Sakmarian, Artinskian, and Kungurian ages.1
  • Guadalupian Epoch: Named for the Guadalupe Mountains in West Texas, it contains the Roadian, Wordian, and Capitanian ages.1
  • Lopingian Epoch: Named for its type area in China, it consists of the Wuchiapingian and Changhsingian ages.1

While these conodont-based zonations are standard for paleoequatorial warm-water provinces, researchers use different zonations for colder regions like Gondwana, where local series and stage names remain necessary for identifying nonmarine successions.1