How does cork cambium contribute to bark formation?

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In the context of plant development, the cork cambium—also known as the phellogen—is a lateral meristem essential for the formation of bark in woody plants.21

As a plant increases in girth through secondary growth, the original epidermis cannot expand and is eventually destroyed. The cork cambium replaces this protective layer by producing cork cells toward the outside and parenchyma cells (phelloderm) toward the inside. Together, these tissues form the [periderm].2

The cork cambium contributes to bark formation through the following processes:

  • Protective Barrier: The cork cells produced by the cambium become nonliving at maturity and are lined with suberin, a fatty, waterproof substance that protects the plant from desiccation and environmental stress.21
  • Continuous Renewal: Because the outer bark cannot accommodate the increasing diameter of the stem, it is shed over time. To maintain protection, new cork cambiums periodically arise in the secondary phloem, creating successive layers of periderm.23
  • Bark Composition: Bark is defined as all tissues outside the vascular cambium. The cork cambium is responsible for generating the outer bark, which consists of these dead cork cells and old, crushed phloem.21