How did Carl Kling find evidence that healthy carriers could spread poliovirus?

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Polio vaccine — polio A health care worker giving a polio vaccine to a child in Katsina state, Nigeria, 2014.

During a major polio epidemic in 1911 that involved 3,840 cases, Carl Kling and his colleagues in Stockholm provided evidence that healthy individuals could transmit the virus. By conducting clinical investigations during this outbreak, they successfully recovered the poliovirus from both paralytic patients and healthy carriers.1

Additionally, in their study of several fatal cases from the same epidemic, Kling and his team identified the presence of the virus within the victims' throats and the tissues of their small intestines.1