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How are statocysts used for balance in invertebrates?

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Meissner's corpuscle; mechanoreception — Mechanoreception In human skin, specialized nerve endings known as Meissner's corpuscles (or tactile corpuscles) are sensitive to touch. They are one of several different types of mechanoreceptors found in human skin.

Statocysts are specialized gravity receptors used by many invertebrates to maintain equilibrium and orient themselves in space. A statocyst typically consists of a fluid-filled vesicle containing one or more dense, sandy, or stonelike elements known as statoliths.1

The mechanism functions as follows:
* Sensory Stimulation: The interior wall of the vesicle is lined with sensory cells equipped with hairs. Because statoliths are heavy, they are pulled downward by gravity. As an animal changes its position, the statolith shifts, either pressing against or shearing across these sensory hairs.12
* Neural Response: This mechanical stimulation triggers sensory cells to send electrical impulses to the animal's nervous system, signaling its orientation relative to gravity.1
* Compensatory Reflexes: Invertebrates use this information to perform compensatory movements—such as adjusting the eyes, head, or limbs—to maintain or restore their original body position.1

While many invertebrates possess these organs, some, such as insects, rely on specialized tactile hairs on their body surface to detect gravity instead. In more advanced invertebrates like cephalopods, statocysts are closed, fluid-filled structures, whereas in primitive forms, they may remain open to the surrounding environment.21