Where do Ophrys orchids attach pollinia on different insect pollinators?

In the genus Ophrys, the placement of pollinia on an insect depends on the specific behavior the orchid induces during pseudocopulation. Because these orchids use sexual deception to attract pollinators, the location of the attachment is determined by where the insect attempts to mate with the flower.12

• Head Attachment: Orchids pollinated by the wasps Trielis and Gorytes, as well as the bee Eucera, induce the insect to attempt copulation with the apex of the flower's lip. In these cases, the pollinarium is affixed to the insect's head.2
• Abdomen Attachment: Orchids pollinated by bees of the genus Andrena generally stimulate the insect to reverse its position and attempt to copulate with the base of the lip. Consequently, the pollinarium is attached to the insect's abdomen.2

It is important to note that this process is entirely deceptive; the male insects receive no reward for their efforts and often expend significant energy in these futile mating attempts.1