What causes the leaf scars found on branches of Magnoliaceae plants?

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The leaf scars found on the branches of Magnoliaceae plants are caused by the shedding of stipules.

Unlike other families within the order Magnoliales, members of the Magnoliaceae family possess these relatively large, leaflike appendages at the base of the leaf stalk. As the leaf expands, the stipules fall off, leaving behind a distinct scar on the branch.1