How does Gaultheria use both wind and animal dispersal?

Most species of Gaultheria, a genus within the Ericales order, utilize a dual-method approach to seed dispersal made possible by their unique fruit structure.1

These plants produce dry, capsular fruits that contain numerous minute seeds, which are adapted for wind dispersal. Simultaneously, these capsules are surrounded by fleshy, brightly colored sepals. This succulent exterior attracts animals—primarily birds, and sometimes mammals—which consume the fleshy parts and disperse the hard seeds through their excrement. By combining a wind-dispersible internal capsule with an animal-attractive external structure, Gaultheria effectively leverages both biotic and abiotic mechanisms to distribute its seeds.21