How do long shoots and spur shoots differ on a ginkgo tree?

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On a Ginkgo tree, the distinction between long shoots and spur (short) shoots is a fundamental aspect of its growth habit and morphology:

  • Growth and Development: Long shoots are responsible for the tree's primary extension, producing leaves during the spring growth period. In contrast, spur shoots are lateral branches that grow much more slowly; after several years, they develop into short, stubby outgrowths from the stem. While the vascular cambium in long shoots produces significant secondary xylem (wood) with well-defined growth rings, the activity in spur shoots is limited, resulting in only a small amount of soft wood each year.12
  • Leaf Production: Leaves on long shoots are typically produced individually during spring growth and are often divided by a deep sinus into two lobes. Spur shoots, however, form clusters of leaves in subsequent years. The degree of leaf lobing also differs; bilobed and undivided leaves are commonly found on spur branches, whereas leaves on the upper parts of long branches are more frequently divided.1
  • Reproduction: Reproductive structures, including both pollen-producing strobili and ovules, are restricted to the spur branches.21