Britannica Sources used in this answer

How do horsetail sporangia release their spores?

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Horsetail (Equisetum) sporangia release their spores through a longitudinal slit on their inner side.1

The dissemination of these spores is facilitated by four ribbon-like appendages known as elaters, which are formed from the outermost wall layer of the spore. These elaters are highly sensitive to moisture; they coil and uncoil in response to changes in humidity, which helps to disperse the spores from the sporangia.21