How do cyanobacteria in coralloid roots benefit cycads?

Cycads benefit from the presence of cyanobacteria within their coralloid roots primarily through the process of nitrogen fixation.12

These symbiotic bacteria convert atmospheric nitrogen into a form that the plant can utilize for its growth and development. Furthermore, in association with the root tissues, these cyanobacteria facilitate the production of beneficial amino acids, specifically asparagine and citrulline, which support the plant's nutritional needs.21