How did the former families Bombacaceae, Tiliaceae, and Sterculiaceae become part of Malvaceae?

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The integration of the former families Bombacaceae, Sterculiaceae, and Tiliaceae into the family Malvaceae is a result of modern plant phylogeny studies, which have largely moved away from older, morphology-based classification systems like that of Arthur Cronquist.1

Since the 1990s, the Angiosperm Phylogeny Group (APG) has utilized molecular techniques—specifically the sequencing of DNA from chloroplasts and nuclei—to establish more robust and testable evolutionary relationships. These molecular analyses revealed that the traditional, morphology-based groupings often failed to represent true monophyletic lineages (groups derived from a single common ancestor).1

Under the current APG system, these formerly segregate families are now included within a broadly defined Malvaceae to ensure the family reflects a single, cohesive evolutionary lineage. This taxonomic revision is part of a broader effort to organize plant diversity based on genetic data, leading to the updated circumscriptions found in the APG IV system.21