Which agriculturalist groups are associated with the Sua, Aka, Efe, and Mbuti peoples?

In the Ituri Forest, the nomadic hunting and gathering peoples—collectively known as the Bambuti—maintain long-standing relationships of mutual interdependence with neighboring village-dwelling agriculturalist groups. Each of the four distinct Pygmy populations is associated with a specific tribe of Bantu- or Sudanic-speaking agriculturalists:12

• Sua: Associated with the Budu (Babudu) people, located on the western edge of the Ituri Forest near Wamba.2
• Aka: Associated with the Mangbetu people, primarily found in the northwest.2
• Efe: Associated with the Sudanic-speaking Mamvu and Lese (Walese) peoples, possessing the broadest distribution across the northern and eastern portions of the forest.2
• Mbuti: Associated with the Bila (Babila) people, living in the central region of the forest.2

These associations are characterized by close economic and ritual ties, often passed down through generations, where the Bambuti trade forest resources like meat and honey for agricultural products and manufactured goods.23