Which rivers did Alexander von Humboldt travel along to reach the Orinoco River?

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To reach the Orinoco River, Alexander von Humboldt and his companion, Aimé Bonpland, traveled south from Caracas through grasslands and scrublands until they reached the banks of the Apure River. From there, they continued their journey by canoe along the Apure until it joined the Orinoco.1