How can flutter tonguing change the sound of a wind instrument?

Flutter tonguing is a specialized performance technique used on wind instruments to alter their timbre, or quality of sound.1

The effect is produced by a rapid, vibratory movement of the tongue, similar to rolling or trilling an "r" while playing. This action creates a distinct, fluttering texture in the tone that differs significantly from standard articulation. While this technique is possible on most wind instruments, it is particularly effective and agile on the flute.21