Which perissodactyls mainly browse on leaves and shoots?

Within the order Perissodactyla, several groups are classified as browsers, meaning they primarily feed on the foliage of trees and shrubs rather than grass.1

• Tapirs: These animals are browsers that inhabit deep forests or swamps. They use their specialized snouts—formed by the fusion of the upper lip and a short proboscis—to pluck leaves and twigs.12
• Rhinoceroses: Most rhinoceros species are browsers. They use their forelimbs to break down trees and shrubs and utilize their lips to reach otherwise inaccessible leaves and twigs. Notably, most rhinoceroses possess a pointed upper lip with a fingerlike process specifically adapted for plucking this vegetation. (The white rhinoceros is a notable exception, as it is a specialized grazer with a broad, square muzzle adapted for feeding on grass.)1
• Extinct Groups: Certain fossil perissodactyls, such as the brontotheres and the anchitheres (a subfamily of early horses), are also identified as browsers.3