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Which animals prey on winged adult mayflies?

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Winged adult mayflies are subject to predation by a variety of animals depending on whether they are in flight or at rest.

When in flight, adult mayflies are consumed by:
* Birds1
* Bats1
* Predatory insects, including dragonflies, robber flies, and hornets1

When at rest, they may be preyed upon by:
* Spiders1
* Beetles1
* Birds1
* Certain mammals, specifically flying squirrels in North America1

Additionally, female mayflies are vulnerable to fishes while they are depositing their eggs on the water's surface.1