Where in puffer fishes is tetrodotoxin especially concentrated?

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In puffer fishes (family Tetraodontidae), the potent neurotoxin known as tetrodotoxin is especially concentrated in the internal organs, also referred to as the viscera.123

Because this substance is highly toxic and can be fatal, the consumption of puffer fish—often prepared as fugu in Japan—requires the fish to be carefully cleaned and prepared by a specially trained chef to ensure the poisonous parts are removed.123