What happens to hippopotamuses during droughts or famines?

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During times of drought or famine, hippopotamuses may be forced to undertake overland migrations, which often result in many deaths.1

Under normal conditions, hippos rely on specific aquatic "day living spaces" to remain submerged, as they lose water rapidly through their skin and require these periodic dips to avoid dehydration and regulate their body temperature. During the dry season, these pools can become extremely crowded, with as many as 150 individuals sharing a single area. When these resources fail due to environmental stress, the animals' survival is severely compromised.21