How do toucans use their serrated bills when feeding?

Toucans use their serrated, or saw-edged, bills to obtain food by tearing open fruits, securing insects, or plucking nestlings from the nests of other birds.12

When feeding on small fruits, toucans often pick them up daintily at the tip of the bill before tossing back their heads to swallow them. While the bill is highly specialized, its exact size and structure remain a subject of study, as it appears larger than what is strictly necessary for these feeding functions alone.12