How do orangutans move through the forest canopy?
Orangutans are primarily arboreal animals, spending more than 90 percent of their time in the trees. To move through the forest canopy, they typically scramble by using all four hands and feet, a method for which they are well-adapted due to their flexible hip joints and hooklike hands and feet with opposable big toes. Because they grip vertical tree trunks equally well with both their forelimbs and hind limbs, they were historically referred to as "four-handed" or quadrumanal.12
While scrambling is their primary mode of travel, orangutans also occasionally swing through the trees using only their arms, a form of locomotion known as brachiation. Although they possess proportionately the longest arms of any great ape, they do not rely on this method as exclusively as some other primates. When on the ground, orangutans are relatively slow and move by walking on their closed fists or extended palms rather than by knuckle-walking like African apes.13