How do male hippopotamuses warn rivals that a territory is occupied?

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To warn rivals that a territory is occupied, male hippopotamuses engage in a specific routine display. Adjacent territorial bulls will stare at each other before turning to expose their rear ends out of the water. They then use their tails to rapidly wag back and forth, flipping feces and urine in a wide arc.1

Additionally, both territorial and subordinate males create dung piles along pathways leading inland, which likely serve as olfactory signposts or scent markers to communicate at night.1