How do leeches swim?

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When swimming, leeches maintain their dorsoventral muscles in a contracted state. This posture allows them to produce undulations of the body through waves of contraction generated by their longitudinal muscles.1

While leeches are well-known for their "contract-anchor-extend" method of crawling along surfaces using their anterior and posterior suckers, their swimming mechanism relies on these rhythmic muscular undulations to propel them through the water.12